Areas of research: physiology of fungal spore germination, morphogenesis and differentiation in fungi, biological control, biology of soilborne pathogens, interactions between soil pathogens and soil biota, soil suppressiveness, integrated control, bacterial diseases, microflora of stored products, degradation of pesticides and pollutants in soil bioremediation, nitrogen fixation, microbial ecology, cellulose degradation, edible mushrooms, water microbiology