physiology of fungal spore germination, morphogenesis and differentiation in fungi, biological control, biology of soilborne pathogens, interactions between soil pathogens and soil biota, soil suppressiveness, integrated control, bacterial diseases, microflora of stored products, degradation of pesticides and pollutants in soil bioremediation, nitrogen fixation, microbial ecology, cellulose degradation, edible mushrooms, water microbiology
Taxonomy of fungi, cereal diseases, diseases of wild grasses, wild cereals as sources for disease resistance, taxonomy of Helminthosporia and related fungi pathogenic to Gramineae, entomophagous fungi as potential biocontrol agents and biocontrol of weeds by pathogens using naturally occurring fungal pathogens
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